Microbial genetics, horizontal gene transfer, mobile
genetic elements, site-specific recombination, molecular
mechanisms of microbial evolution.

1958~1959 University of Southern California, Los Angeles,
U.S.A. 연구원
1960~1970 University of Geneva 연구원, 조교수, 부교수
1961~1972 Swiss Committee of the International Union for
Pure
and Applied Biophysics 회장
1967~1978 Current Topics of Microbiology and Immunology 편
집위원
1967~ Molecular and General Genetics 편집위원
1968~1975 Quarterly Reviews of Biophysics 편집위원
1970~1971 University of California, Berkeley, USA 방문교수
1971~1996 University of Basel 교수
1981~1985 The EMBO Journal 편집위원
1981~ European Academy of Arts, Sciences and Humanities
회원
1984~ National Academy of Sciences, USA 외국인준회원
1984~1985 University of Basel, Faculty of Science 학장,
Rector
1986~1988 University of Basel, Rector
1986~ Molecular Microbiology 편집위원
1988~ Swiss Academy of Engineering Sciences 회원
1989~ Academia Europaea 회원
1996~1999 President, International Council for Science

1962 University of Geneva, Prix Plantamour-Pr
vost
1978 노벨의학생리학상